The Firefly would be deployed as one tank per troop of Cromwell or Sherman tanks. The deployment with Cromwell troops made servicing and supply of those units more complex. The Firefly was also slower than the Cromwell.
While the number of Panthers and Tigers only accounted for some 30% of the nearly 2,500 German tanks deployed in Normandy (the rest being composed of Panzer IVs, Sturmgeschütz IIIs and other tanks the standard Shermans were able to effectively handle), the importance of Caen to the Germans and Montgomery's operations which pinned German armored forces in front of the British positions so the American units could break out to the west meant that British and Commonwealth units had to face over 70% of all German armour deployed during the Battle of Normandy, as well as almost all the elite, well-equipped SS units which contained the fearsome Tigers and Panthers. Thus, despite the relatively low number of Panthers and Tigers deployed, they would almost all be facing British and Commonwealth troops. As a result, the Sherman Firefly was perhaps the most valued tank by British and Commonwealth commanders, as it was the only tank in the British Army able to effectively defeat the Panthers and Tigers at the standard combat ranges in Normandy.
This fact did not go unnoticed by the Germans, who realized that these long-barrel Shermans posed a much greater threat to their heavy tanks than the regular Shermans, and German tank crews and anti-tank gun crews were instructed to eliminate Fireflies first. Similarly, the Firefly crews realized that the distinctive long barrel of their 17 pounder gun made the Firefly stand out from regular Shermans, so crews attempted to disguise their tanks in the hope they would not be targeted. Some crews had the front half of the gun barrel painted white on the bottom and dark green or the original olive drab on the top to give the illusion of a shorter gun barrel. Another suggestion was for a shorter wooden dummy gun would be mounted on the rear of the turret and point forward; however, this tactic does not appear to have been used in combat.
Despite this, the Fireflies increased firepower was much valued by British and Commonwealth tankers, and during many engagements, the Firefly proved its worth, knocking out the feared Tigers and Panthers at long range, as well as less formidable tanks like the Mark IVs and StuGs.
One example of this increased firepower was displayed by Lt. G.K Henry's Firefly during the defense of Norrey-en-Bessin on 9 June against an attack by the 3rd Company of the 12th SS Panzer Regiment of the 12th SS Panzer Division. Determined to capture the town in preparation for a larger offensive to drive the British and Canadians back into the sea, Kurt Meyer ordered an attack by 12 Panthers of the 3rd Company and infantry to attack Norrey-en-Bessin and drive the Canadians out of the town. The attack got underway at 1300 hours with the Panthers racing to the town at full speed only to stop to fire their guns, quickly outrunning their infantry support which was forced to ground by allied artillery fire. Within 1,000 m (1,100 yd) of the town, 9 Shermans of the 1st Hussars opened fire into the advancing Panthers flanks. Lt. Henry's gunner, Trooper A. Chapman, waited until the Panthers "lined up like ducks in a row" and quickly knocked out five German Panthers with just 6 rounds. The attack was repulsed with the loss of 7 of the 12 attacking Panthers, the majority credited to Lt. Henry's single Firefly.
Another similar example occurred on 14 June, during Operation Perch, Sgt. Harris of the 4th/7th Dragoon Guards, along with three standard Shermans, set up defensive positions along with the infantry after successfully driving out the Germans in the village of Lingèvres, near Tilly-sur-Seulles. Looking through his binoculars, Sgt. Harris spotted two Panthers advancing from the east. He opened fire at a range of 800 metres (870 yd), knocking out the lead Panther with his first shot, and the second Panther with his second. Relocating to a new position on the other side of the town, he spotted another three Panthers approaching from the west. From his well-concealed flanking position, he and his gunner Trooper Mackillop eliminated all three with just three rounds. Harris and his gunner had knocked out five Panthers with as many rounds, once again demonstrating the potency of the Firefly, especially when firing from a defensive position on advancing enemy tanks.
Tanks of the 1st Northamptonshire Yeomanry and elements of the 51st (Highland) Division reached the French village of Saint-Aignan-de-Cramesnil on the morning of 8 August 1944 during Operation Totalize. While B Squadron stayed around the village, A and C Squadrons moved further south into a wood called Delle de la Roque.C Squadron positioned themselves on the east side of the woods and the understrength A Squadron in the southern portion with No. 3 Troop on the western edge of the wood. From this position, they overlooked a large open section of ground and were able to watch as German tanks advanced up Route nationale 158 from the town of Cintheaux. Under strict orders from the troop commander, they held their fire until the German tanks were well within range. Ekins, the gunner of Sergeant Gordon's Sherman Firefly (called Velikye Luki - A Squadrons tanks were named after towns in the Soviet Union) had yet to fire his gun in action. With the Tiger tanks in range, the order was given to fire, what followed was an almost twelve minute battle that saw Ekins destroying all three Tigers that No. 3 Troop could see (there were actually 7 Tiger tanks in the area heading north along with some other tanks and self propelled guns). A short time later, the main German counterattack was made in the direction of C Squadron. A Squadron (less Sgt Gordon who had been wounded and had already bailed out of the Firefly) moved over to support them and in the resulting combat, Ekins destroyed a Panzer IV before his tank was hit and the crew were forced to bail out. One of the Tigers Ekins is credited with knocking out was that of Michael Wittmann, though there is still some controversy over whether Ekins really killed Wittman as Fireflies of the Sherbrooke Fuisilier Regiment were also firing at the Tigers from a closer range of 150 m (160 yd).
Overall the Firefly proved itself a very successful tank despite the fact it was only intended as a stopgap tank until future British tanks like the Comet and the Centurion came into service. While Normandy had priority, Fireflies also served with distinction in Italy in British and Commonwealth units.
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